Overall, there were three interviews:
1) Within a week of applying, I had a 30 minute screening interview with a recruiter (via phone). They asked about some points on my CV/application, and what my timeline for starting the position would be.
2) The next day, I had a 30 minute video interview with the Chief Editor, who asked more about my research experience and interest in the position, as well as some questions to gauge my familiarity with the publishing/review process.
3) The next week, I had a ~4 hour video interview and manuscript test. They emailed me three manuscripts (molecular biology, neuroimaging, and animal models), and gave an hour to review/critique them, before meeting with the Chief Editor and an Associate Editor for an hour to discuss the strengths/weaknesses of each paper, and whether or not I would send it out to review. The rest of the interview consisted of more questions about my background/motivation from the Chief Editor and a Senior Editor.
Two weeks after the manuscript test, I received an informal offer via email. Overall the editors were very friendly, it was just difficult to gauge how I was doing at each stage of the interview.